This is the heart of the game, and the single most misexplained mechanic in competing reviews. Read the reel restrictions carefully, because they define exactly what you are hoping to see.
Trigger condition. The Rush Express feature activates when a Gold Bell or Gold Train symbol lands on reel 5, and at least one Golden Nugget or Coloured Train symbol lands on reels 1 to 4 on the same spin. Nuggets alone do nothing. A collector alone does nothing. You need both halves of the equation, and they cannot appear on each other's reels.
Mode 1, Rush Express (Bell). When the Bell lands on reel 5, every visible nugget's cash value is added together and paid instantly. Clean, immediate, no multiplier.
Mode 2, Rush Express (Gold Train). When the Gold Train lands on reel 5, all visible nugget values are collected onto the train above the reels. The train then awards that combined total up to 10 times, applying a random multiplier of 1x to 10x to the whole collected sum. This is the mode that produces the game's headline screenshots.
Coloured Trains. Coloured Train symbols land on reels 1 to 4 in four colours matching the four jackpot tiers. When a Rush Express feature triggers, each Coloured Train runs its own mini-feature: a train of carriages rolls past, each carriage adding a cash prize to that symbol's total. The final carriage can award a random multiplier, or the jackpot of the corresponding colour. That is a second, independent route into the jackpot ladder.
The strategic implication. Because the Train mode can multiply the entire collected sum, the value of a nugget-heavy screen scales dramatically. A screen holding A$12 of nuggets is worth A$12 with the Bell and up to A$120 with the Train. There is nothing a player can do to influence which collector lands. Understanding the difference simply explains why two visually similar wins can differ tenfold, and why chat rooms fill with screenshots of the Train and almost none of the Bell.